Natural Bridge ZooRICHMOND, Va. (AP) _ The Humane Society of the United States says a months-long undercover investigation at the private Natural Bridge Zoo found abuse and neglect leading to the deaths of a giraffe and other animals. The Humane Society outlined the allegations Thursday, focusing as well on what it called the persistent mistreatment of tiger cubs. The society says the cubs were hit and punched starting when they were 6 weeks old. The Humane Society investigation also found a critical lack of veterinary care, crowded conditions and filthy cages, among other unsafe conditions. The society said it has filed complaints with state, local and federal officials. In a written response, Natural Bridge Zoo owner Karl Mogensen called the allegations “slanderous” and “vicious propaganda” aimed at soliciting donations. He also denied the accusations.